生物专业英语第三版蒋悟生编__课文翻译(7)

中心，其直接参与光合作用。大部分光反应细胞器拥有两套反应中心，P680和P700，每个光系统都含有一个电子受体和电子供体。这些集合体就是大家熟识的光合系统Ⅰ和光合系统Ⅱ 。

The Light-Dependent Reaction: Converting Solar Energy into Chemical-Bond Energy

光反应：光能转化成化学键能

The photosystems of the light-dependent reactions are responsible for the packaging of light energy in the chemical compounds ATP and NADPH. This packaging takes place through a series of oxidation reduction reactions set in motion when light strikes the P680 reaction center in photosystem Ⅱ. In this initial event water molecules are cleaved, oxygen is released, and electrons are donated. These electrons are accepted first by plastoquinone and then by a series of carriers as they descend an electron transport chain. For each four electrons that pass down the chain, two ATPs are formed. The last acceptor in the chain is the P700 reaction center of photosystem Ⅰ. At this point incoming photons boost the energy of the electrons, and they are accepted by ferredoxin. Ferredoxin is then reoxidized, and the coenzyme NADP+ is reduced to the NADPH. The ATP generated previously and the NADPH then take part in the light independent reactions.

光反应的光系统将光能转化成化学复合物ATP和NADPH。当光激活光系统Ⅱ的光反应中心时，通过一系列的氧化还原反应实现能量的传递。反应开始时，水被分解，氧被释放并提供电子。电子首先传递给质体醌，然后通过一系列载体形成的电子传递链。每传递4个电子，形成2个ATP。最后一个受体存在于光反应系统Ⅰ的反应中心里。此处光子激活电子，电子传递给铁氧还蛋白。铁氧还蛋白再氧化，并且辅酶NADP+还原成NADPH。早期产生的ATP和NADPH进入暗反应。

The production of ATP from the transport of electrons excited by light energy down an electron transport chain is termed photophosphorylation. The one-way flow of electrons through photosystems II and I is called noncyclic photophosphorylation; plants also derive additional ATP through cyclic photophosphorylation, in which some electrons are shunted back through the electron transport chain between photosystems Ⅱ and Ⅰ.

由电子传递链偶连产生ATP的过程称为光合磷酸化。通过光合系统Ⅱ流经光合系统Ⅰ的电子路径称非循环式光合磷酸化；植物通过循环式光合磷酸化获得额外的ATP，一些电子在光合系统Ⅰ和Ⅱ之间的电子传递链中回流。

The Light-Independent Reactions: Building Carbohydrates

暗反应：碳水化合物的形成

In the light-independent reactions of photosynthesis, which are driven by ATP and NADPH, C02 is converted to carbohydrate. The reactions are also known as the Calvin-Benson cycle. Atmospheric CO2, is fixed as it reacts with ribulose biphosphate (RuBP), a reaction that is catalyzed by the enzyme ribulose biphosphate carboxylase. The reduction Of C02 to carbohydrate (fructose diphosphate) is completed via several more steps of the cycle. Finally, RUBP is regenerated so that the cycle may continue.

由ATP和NADPH驱动的暗反应中，二氧化碳转化成碳水化合物。即卡尔文循环。二磷酸核酮糖固定二氧化碳，由二磷酸核酮糖羧化酶催化。

Oxygen: An Inhibitor of photosynthesis（氧：光合作用的抑制因子）

High levels of oxygen in plant cells can disrupt photosynthesis and can also cause photorespiration-an inefficient fun of the dark reactions in which 02 is fixed rather than C02 and no carbohydrate is produced.

Reprieve from Photorespiration: The C4 Pathway

Most plants are C3 plants; they experience decreased carbohydrate production under hot, dry conditions as a result of the effects of photorespiration. Among C4 plants, however, special leaf anatomy and a unique biochemical pathway enable the plant to thrive in and conditions. Thus C4 plants lessen photorespiration by carrying out photosynthesis only in cells that are insulated from high levels of CO2. They also possess a novel mechanism for carbon fixation.

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